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Additionally, the study detected a total of 106 O -glycosylation sites in CSF proteins, and they predominantly occurred on Ser or Thr residues that had a Pro residue at one of the three positions: P-S/T, S/T-P, and S/T-X-X-P, pointing toward a potential consensus motif for O -glycosylation that warrants further investigations (Halim et al., 2013)